1. Department of Cancer Biology, University of Pennsylvania, Philadelphia, PA 19104, USA
2. Cell and Molecular Biology Graduate Group, University of Pennsylvania, Philadelphia, PA 19104, USA
3. Department of Radiation Oncology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA 19104, USA
4. Department of Biomedical Sciences, University of Pennsylvania, School of Veterinary Medicine, Philadelphia, PA 19104, USA
5. Abramson Family Cancer Research Institute, Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA 19104, USA