1. Wellcome Trust Sanger Institute, Hinxton, Cambridge CB10 1SA, UK
2. Current address: Wellcome Trust–Medical Research Council Cambridge Stem Cell Institute, University of Cambridge, Tennis Court Road, Cambridge, CB2 1QR, UK
3. Centre for Modeling Human Disease, Toronto Centre for Phenogenomics, 25 Orde Street, Toronto, M5T 3H7, Canada
4. Current address: National Institute for Medical Research, The Ridgeway, Mill Hill, London, NW7 1AA, UK
5. Current address: LabPMM GmbH, Bunsenstr. 7a, 82152 Martinsried, Germany
6. Current address: Wolfson Centre for Age-Related Diseases, King’s College London, London, SE1 1UL, UK