1. Department of Biology, St. Mary's College of Maryland 1 , St. Mary's City, MD 20686 , USA
2. Miami University 2 Department of Biology and Center for Visual Sciences , , Oxford, OH 45056 , USA
3. School of Molecular & Cellular Biology, University of Illinois at Urbana-Champaign 3 , Urbana, IL 61801 , USA
4. Research Institute of Molecular Pathology, Vienna Biocenter 4 , 1030 Vienna , Austria