1. Department of Biology, University of North Carolina at Chapel Hill, Chapel Hill, NC 27599-3280, USA
2. Lineberger Comprehensive Cancer Center, University of North Carolina at Chapel Hill, Chapel Hill, NC 27599-3280, USA
3. Department of Biological Sciences, Carnegie Mellon University, Pittsburgh PA 15213, USA
4. Curriculum in Genetics and Molecular Biology, University of North Carolina at Chapel Hill, Chapel Hill, NC 27599-3280, USA