1. Integrated Program in Cellular, Molecular, and Biophysical Studies, Columbia University College of Physicians and Surgeons, New York, NY 10032, USA
2. Department of Pathology, Columbia University College of Physicians and Surgeons, New York, NY 10032, USA
3. Department of Anatomy and Cell Biology, Columbia University College of Physicians and Surgeons, New York, NY 10032, USA