1. The Salk Institute for Biological Studies, PO Box 85800, San Diego, CA 92186,USA
2. Present address: McGill University Centre for Research in Neuroscience, 1650 Cedar Avenue, Montreal, QC H3G 1A4, Canada
3. Department of Biology, University of California, Riverside, CA 92521,USA
4. Present address: CyThera, San Diego, CA 92121, USA