1. Wellcome Trust – Medical Research Council Stem Cell Institute, University of Cambridge 1 , Cambridge CB2 0AW , UK
2. Randall Centre for Cell and Molecular Biology, King's College London 2 , London SE1 1YR , UK
3. Altos Labs Cambridge Institute of Science, Granta Park 3 , Cambridge CB21 6GP , UK
4. Institute of Cell Biology and Neuroscience, Goethe University, 4 Frankfurt 60439 , Germany
5. Institute of Bioengineering, School of Life Sciences, Ecole Polytechnique Fédérale de Lausanne (EPFL) 5 , Lausanne CH-1015 , Switzerland
6. Living Systems Institute, University of Exeter 6 , Exeter EX4 4QD , UK