1. Oxford Brookes University 1 Department of Biological and Medical Sciences , , Gipsy Lane, Oxford OX3 0BP , UK
2. , University of Oxford 2 Peter Medawar Building for Pathogen Research , Oxford OX1 3SY , UK
3. 3 Biology Centre CAS, Institute of Parasitology, Branišovská 1160/31, 370 05 České Budějovice, Czech Republic