1. Faculty of Environment, Science and Economy, University of Exeter, Exeter EX4 4QJ, United Kingdom
2. Marine Spatial Ecology Lab, School of the Environment The University of Queensland, St Lucia, QLD 4072, Australia
3. National Oceanic and Atmospheric Administration, Atlantic Oceanographic and Meteorological Laboratory, Ocean Chemistry and Ecosystem Divisions, Miami, FL 33149
4. Commonwealth Scientific and Industrial Research Organisation, Oceans & Atmosphere, St Lucia, QLD 4000, Australia