1. Ecology and Evolutionary Biology Department, Yale University, New Haven, CT 06511
2. Centre for African Ecology, School of Animal, Plant, and Environmental Sciences, University of the Witwatersrand, Johannesburg 2000, South Africa
3. Biological Sciences Department, University of Cape Town, Cape Town 7700, South Africa
4. Scientific Services, Kruger National Park, South African National Parks, Skukuza, Private Bag x 402, South Africa
5. School of Natural Resource Management, Nelson Mandela University, George 6530, South Africa
6. Senckenberg Biodiversity and Climate Research Centre, 60325 Frankfurt am Main, Germany
7. Conservation Management, Kruger National Park, South African National Parks, Skukuza, Private Bag x 402, South Africa
8. Agence Nationale des Parcs Nationaux, Libreville, BP 20379, Gabon
9. Soil, Crop and Climate Sciences Department, University of the Free State, Bloemfontein 9300, South Africa
10. College of Biological Sciences, University of Minnesota, St. Paul, MN 55108
11. Department of Forest Resources, University of Minnesota, St. Paul, MN 55108