1. Nordcee, Department of Biology, University of Southern Denmark, Odense M 5230, Denmark
2. Center for Ocean Life, National Institute of Aquatic Resources, Technical University of Denmark, Kongens Lyngby, 2800 Denmark
3. Danish Institute of Advanced Studies, Odense M, 5230 Denmark
4. Key Laboratory of Petroleum Geochemistry, Research Institute of Petroleum Exploration and Development, China National Petroleum Corporation (Petrochina), Beijing, 100093 China