1. Department of Zoology, University of Oxford, Oxford, OX1 3PS, UK
2. Central Research Institute of Electric Power Industry, 1646, Abiko, Abiko, Chiba 270–1194, Japan
3. Graduate School of Environmental Studies, Nagoya University, Furo-cho, Chikusa-ku, Nagoya 464–8601, Japan
4. College of Agriculture and Life Sciences, Cornell University, 177 Roberts Hall, Ithaca, NY 14853, USA
5. Department of Fisheries Sciences, Hokkaido University, Hakodate, Japan