1. Horned beetles;W.G. Eberhard,1980
2. Diversity in the weapons of sexual selection: horn evolution in the beetle genus Onthophagus (coleoptera: scarabaeidae).;D.J. Emlen,2007
3. Conservation, innovation, and bias: Embryonic segment boundaries position posterior, but not anterior, head horns in adult beetles;H.A. Busey,2016
4. Structural adaptations to diverse fighting styles in sexually selected weapons;E.L. McCullough,2014
5. Mate securing tactics and the cost of fighting in the Japanese horned beetle, Allomyrina dichotoma L. (Scarabaeidae);M.T. Siva-Jothy,1987