1. These authors contributed equally
2. Université de Paris, Paris, France
3. Evolutionary genomics of RNA viruses, Institut Pasteur, Paris, France
4. Molecular Genetics of RNA Viruses, CNRS - UMR 3569, University of Paris, Institut Pasteur, Paris, France
5. National Reference Center for Respiratory Viruses, Institut Pasteur, Paris, France
6. Mutualized Platform of Microbiology, Pasteur International Bioresources Network, Institut Pasteur, Paris, France
7. National Influenza Centre, Viral Respiratory Laboratory, Algiers, Algeria
8. These authors co-supervised this work