1. College of Life Sciences, Ritsumeikan University, Kusatsu, Shiga 525-8577, Japan
2. Graduate School of Science, Osaka University, Toyonaka, Osaka 560-0043, Japan
3. Graduate School of Environmental and Life Science, Okayama University, Okayama 700-8530, Japan
4. Graduate School of Agricultural and Life Sciences, The University of Tokyo, Bunkyo, Tokyo 113-8657, Japan