1. Department of Biochemistry and Genome Biology, Hirosaki University Graduate School of Medicine, Hirosaki 036-8562, Japan
2. Department of Biomedical Science, Hirosaki University Graduate School of Health Science, Hirosaki 036-8564, Japan
3. Department of Molecular Function and Physiology, Faculty of Pharmacy, Keio University, Tokyo 105-8512, Japan
4. Department of Ophthalmology, Hirosaki University Graduate School of Medicine, Hirosaki 036-8562, Japan