1. Department of Ophthalmology and Columbia University, New York, New York, United States
2. Schepens Eye Research Institute and Department of Ophthalmology, Harvard Medical School, Boston, Massachusetts, United States
3. Department of Ophthalmology and Columbia University, New York, New York, United States 3Department of Pathology and Cell Biology, Columbia University, New York, New York, United States
4. Department of Ophthalmology, New York University School of Medicine, New York, New York, United States