1. Oulu Center for Cell-Matrix Research, Department of Medical Biochemistry and Molecular Biology, University of Oulu, Oulu, Finland 2Biocenter Oulu, Oulu, Finland
2. Oulu Center for Cell-Matrix Research, Department of Medical Biochemistry and Molecular Biology, University of Oulu, Oulu, Finland
3. Department of Ophthalmology, Oulu University Hospital, University of Oulu, Oulu, Finland
4. Biocenter Oulu, Oulu, Finland 4Departments of Pathology, University of Oulu and Oulu University Hospital, Oulu, Finland
5. Departments of Ophthalmology and Anatomy, Institute for Human Genetics, University of California, San Francisco, School of Medicine, San Francisco, California