1. Rikkyo University 1 Department of Life Science, College of Science, , Tokyo, Japan
2. Keio University School of Medicine 2 Department of Molecular Biology, , Tokyo, Japan
3. National Institute of Genetics 3 Department of Chromosome Science, , Mishima, Japan
4. RIKEN Center for Integrative Medical Sciences 4 Laboratory for Functional Non-Coding Genomics, , Yokohama, Japan
5. Japan Science and Technology Agency (JST), Precursory Research for Embryonic Science and Technology (PRESTO) 5 , Saitama, Japan