1. Division of Life Science, College of Natural Sciences, Gyeongsang National University, Jinju‑si, Gyeongsang 52828, Republic of Korea
2. Natural Medicine Research Center, Korea Research Institute of Bioscience and Biotechnology, Cheonju‑si, Chungcheongbuk‑do 28116, Republic of Korea
3. Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, Cau Giay, Hanoi 01211, Vietnam
4. Department of Biochemistry (BK21 Four), College of Natural Sciences, Kangwon National University, Chuncheon, Gangwon 24414, Republic of Korea