1. Division of Molecular Genetics Research, Life Science Research Center, University of Toyama, Toyama 930‑0194, Japan
2. Department of Ophthalmology, Graduate School of Medicine and Pharmaceutical Sciences, University of Toyama, Toyama 930‑0194, Japan
3. Department of Liberal Arts and Sciences, Toyama Prefectural University, Toyama 939‑0398, Japan