1. Forest Research Department, Gyeongsangnam-do Forest Environment Research Institute, Jinju, 52615, Korea
2. Institute of Agriculture and Life Science, Gyeongsang National University, Jinju, Gyeongnam, 52828, South Korea
3. Division of Environmental Forest Science, Gyeongsang National University, Jinju, 52828, Republic of Korea
4. Department of Medicinal Plant Resources, Nambu University, Wealkye-dong, Gwangsan-gu, Gwangju 506706, Korea