1. Division of Applied Bioseienee, Graduate School of Agriculture, Hokkaido University, North-9 West-9, Kita-ku, Sapporo 060-8589, Japan
2. Faculty of Applied Biological Science, Hiroshima University, Kagamiyama 1-4-4, Higashi- Hiroshima 739-8528, Japan
3. Division of Applied Bioseienee, Graduate School of Agriculture, Hokkaido University, North-9 West-9, Kita-ku, Sapporo 060-8589, Japan / CREST, Japan Science and Technology Corporation (JST), Kawaguchi 332-0012, Japan
4. National Research Institute of Vegetables, Ornamental Plants and Tea, Mii-machi 1823, Kurume 839-8503, Japan