1. Department of Biological Sciences and Biotechnology, Botswana International University of Science and Technology, Palapye, Botswana
2. Department of Zoology and Entomology, Rhodes University, Makhanda, South Africa
3. Aquatic Systems Research Group, Department of Ecology and Resource Management, University of Venda, Thohoyandou, South Africa
4. GEOMAR, Helmholtz-Zentrum für Ozeanforschung Kiel, Kiel, Germany