1. Department of Biological and Environmental Science, University of Jyväskylä, Jyväskylä, Finland
2. Department of Environmental and Biological Sciences, University of Eastern Finland, Kuopio, Finland
3. Oregon Institute of Marine Biology, University of Oregon, Charleston, OR, USA
4. Faculty of Biological and Environmental Sciences, University of Helsinki, Lahti, Finland
5. Lammi Biological Station, University of Helsinki, Lammi, Finland