1. Department of Biology, College of Natural and Computational Sciences, University of Gondar, Ethiopia
2. Department of Zoological Sciences, College of Natural Sciences, Addis Ababa University, Ethiopia
3. Department of Fisheries and Aquatic Sciences, School of Fisheries and Wildlife, College of Agriculture and Environmental Sciences, Bahir Dar University, Ethiopia
4. School of Environmental and Life Sciences, University of Newcastle, Australia; Okavango Research Institute, University of Botswana