1. Biotechnology and Environmental Engineering Team, Laboratory for Process Environmental and Energy Engineering, National School of Applied Sciences, Ibn Zohr University, Agadir, Morocco;
2. Laboratory of Biotechnologies and Valorization of Natural Resources, Faculty of Sciences - Agadir, Ibn Zohr University, Agadir, Morocco;
3. Integrated Crop Production Unit, Institut National de la Recherche Agronomique d’Agadir, Inezgane, Agadir, Morocco;
4. Univ. Lille, INRA, ISA, Univ. Artois, Univ. Littoral Côte d'Opale, EA 7394 - ICV - Institut Charles Viollette, Lille, France;
5. Laboratory of Biology and Biotechnology of Microorganisms, Department of Biology, Faculty of Sciences Semlalia, Marrakech, Morocco & Agrobiosciences Program, University Mohammed IV Polytechnic (UM6P), Benguerir, Morocco;
6. Wheat Health, Genetics and Quality Research Unit, USDA-ARS, Pullman, Washington, USA