1. University of Rwanda, College of Education, School of Education, Department of Mathematics, Science and Physical Education, Rwanda
2. University of Rwanda, College of Science and Technology, Center of Excellence in Biodiversity and Natural Resources Management, Rwanda
3. University of Rwanda, College of Agriculture, Veterinary and Animal Sciences, School of Forestry and Biodiversity Conservation, Rwanda
4. Royal Belgian Institute of Natural Science, Brussels, Belgium
5. Université de Liège, Gembloux Agro Bio-Tech, Entomologie Fonctionelle et Evolutive, Belgique