1. a National Institute of Agro-Environmental Sciences , Ibaraki, Tsukuba , 305 , Japan
2. b CREAF (Center for Ecological Research and Forestry Applications) , Universitat Autònoma de Barcelona , Bellaterra, 08193 , (Barcelona)
3. c IBMBCSIC (Consejo Superior de Investigaciones Científicas) , C/ Jordi Girona 18-26, 08034 , Barcelona , Spain