1. a Marine & Coastal Development Unit , Argyll & Bute Council , Lorn House, Albany Street, Oban, Argyll, PA34 4AR, UK
2. b Smithsonian Tropical Research Institute , Box 0843‐03092, Panama
3. c Heriot‐Watt University , School of Life Sciences , Edinburgh, EH14 4AS, UK
4. d Heriot‐Watt University , School of the Built Environment , Edinburgh, EH14 4AS, UK