1. Nuffield Laboratory of Ophthalmology, Nuffield Department of Clinical Neurosciences, University of Oxford, Oxford, UK
2. Ophthalmic Genetics and Visual Function Branch, National Eye Institute, National Institutes of Health, Bethesda, MD, USA
3. Save Sight Institute, Discipline of Ophthalmology, University of Sydney, Sydney, Australia
4. Vision and Eye Research Institute, Anglia Ruskin University Medical School, Cambridge, UK