1. Institute of Marine Science, University of Auckland, Leigh, New Zealand;
2. Institut de Recherche pour le Development (IRD), UMR-241 EIO, Labex corail, Papeete, Tahiti, Polynésie Française;
3. Université de la Polynésie Française, UMR-241 EIO, Labex corail, Tahiti, Polynésie Française;
4. Aquatic Ecology and Water Quality Management Group, Wageningen University & Research, Wageningen, The Netherlands