1. Department of Entomology and Plant Pathology, North Carolina State University, Raleigh, North Carolina
2. Farlow Herbarium, Department of Organismic and Evolutionary Biology, Harvard University, 22 Divinity Avenue, Cambridge, Massachusetts 02138
3. Department of Geobotany and Botanical Garden, Herbarium, Institute of Biology, Martin Luther University, Neuwerk 21, Halle (Saale) 06099, Germany
4. Departamento de Botánica, Ecología y Fisiología Vegetal, Universidad de La Laguna, Avenida Astrofísico Francisco Sánchez, s/n, Facultad de Farmacia, Apartado 456, Código postal 38200, San Cristóbal de La Laguna, S/C de Tenerife, Canary Islands, Spain