1. Institut de Systématique, Évolution, Biodiversité (ISYEB), Muséum national d’Histoire naturelle, CNRS, Sorbonne Université, EPHE, CP 50, 57 Rue Cuvier, F–75005, Paris, France
2. Sichuan Cave Exploration Team, 610000, Chengdu, PR China
3. Department of Entomology, College of Agriculture, South China Agricultural University, 510642, Wushan, Guangzhou, PR China