1. Biology Department, University of St. Thomas, Saint Paul, MN, 55105, USA;
2. School of Public Health, University of Minnesota, 420 Delaware St. SE, Minneapolis, MN, 55455, USA;
3. Department of Ecology, Evolution, and Behavior, University of Minnesota, 1987 Upper Buford Circle, Falcon Heights, MN, 55108, USA;
4. St. Anthony Falls Laboratory, University of Minnesota, 2 Third Avenue SE, Minneapolis, MN, 55414, USA