1. Biotechnology and Plant Improvement Laboratory, Centre of Biotechnology of Sfax (CBS), University of Sfax, Sfax, Tunisia
2. Plant Production Department, College of Food and Agricultural Sciences, King Saud University, Riyadh, Saudi Arabia
3. Departments of Life Sciences, Faculty of Sciences of Gafsa, Gafsa, Tunisia
4. Laboratoire de Biotechnologie Moléculaire des Eucaryotes, Centre de Biotechnologie de Sfax, Université de Sfax, Sfax, Tunisia