1. Department of Biology, Faculty of Science and Education, Busitema University, Tororo, Uganda
2. Fish Habitat Management Program, National Fisheries Resources Research Institute, National Agricultural Research Organisation, Jinja, Uganda
3. Consultant Environment and Natural Resources, Kampala, Uganda
4. Department of Zoology, Entomology and Fisheries Sciences, College of Natural Sciences, Makerere University, Kampala, Uganda