1. LITTORAL, Ifremer, F‐29900 Concarneau, France
2. Department of Botany and Plant Pathology, Oregon State University, College of Agricultural Sciences, 2082 Cordley Hall, Corvallis, OR 97331-2902, USA
3. Station Biologique de Roscoff, Sorbonne Université, Université Pierre et Marie Curie (Paris VI), Place Georges Teissier, CS90074, 29688 Roscoff, France
4. Station Biologique de Roscoff, CNRS, UMR 7144, Place Georges Teissier, CS90074, 29688 Roscoff, France
5. Sorbonne Université, CNRS, Station Biologique de Roscoff, AD2M, UMR 7144, ECOMAP, Roscoff, France
6. Roscoff Culture Collection, Sorbonne Université, CNRS, FR2424, Station Biologique, Roscoff, 29680, France
7. Graduate School of Bioresources, Mie University, Tsu, Mie 514–8507, Japan
8. National Museum of Natural History, DGD-REVE, Station de Biologie Marine de Concarneau, Place de la Croix, 29900 Concarneau, France