1. Department of Agricultural Sciences, Faculty of Agriculture Natural Resources and Environment, Naresuan University, Phitsanulok 65000, Thailand
2. Center of Excellence in Research in Agricultural Biotechnology, Naresuan University, Phitsanulok 65000, Thailand
3. Walai Rukhavej Botanical Research Institute, and Center of Excellence in Biodiversity Research, Mahasarakham University, Maha Sarakham 44150, Thailand