1. Research Center for Inland Seas, Kobe University, Rokkodai, Kobe 657-8501, Japan
2. Phycology Research Group and Center for Molecular Phylogenetics and Evolution, Ghent University, Krijgslaan 281 (S8), Ghent B-9000, Belgium
3. Marine Laboratory, University of Guam, 303 University Drive, UOG Station, Mangilao 96913, Guam
4. School of Life Sciences, University of Hawaii, 3190 Maile Way, Honolulu, Hawaii 96822, USA
5. UMR-EIO, LabEx-CORAIL, Université de la Polynésie Française, 98702 Faa’a, Tahiti BP 6570, French Polynesia