1. School of History and Geography, Minnan Normal University, Zhangzhou, China
2. State Key Laboratory for Subtropical Mountain Ecology of the Ministry of Science and Technology and Fujian Province, Fujian Normal University, Fuzhou, China
3. School of Geographical Sciences, Fujian Normal University, Fuzhou, China
4. Department of Dendrology and Floriculture, Faculty of Horticultural, Hungarian University of Agriculture and Life Sciences, Budapest, Hungary