1. Laboratory of Anatomy, Department of Basic Veterinary Sciences, Hokkaido University, Sapporo, Japan
2. Laboratory of Agrobiomedical Science, Faculty of Agriculture, Hokkaido University, Sapporo, Japan
3. One Health Research Center, Hokkaido University, Sapporo, Japan
4. Laboratory of Laboratory Animal Science and Medicine, Department of Applied Veterinary Sciences, Hokkaido Universityty, Sapporo, Japan