1. Department of Soil Science, Stellenbosch University, Stellenbosch, South Africa
2. Department of Microbiology, Stellenbosch University, Stellenbosch University Water Institute, Stellenbosch, South Africa
3. Centre for Microbial Ecology and Genomics, Department of Biochemistry, Genetics and Microbiology, University of Pretoria, Genomics Research Institute, Pretoria, South Africa
4. Department of Microbiology, Stellenbosch University, Stellenbosch, South Africa
5. Department of Earth Sciences, Kent State University, Kent, Ohio, USA