1. Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China
2. Yantai Institute of Coastal Zone Research, Chinese Academy of Sciences, Yantai, China
3. Laboratory of Marine Biology and Biotechnology, Qingdao National Laboratory for Marine Science and Technology, Qingdao, China
4. Center for Ocean Mega-Science, Chinese Academy of Sciences, Qingdao, China