1. Centre for Ecological Genomics and Wildlife Conservation, Department of Zoology, University of Johannesburg, Auckland Park, South Africa
2. Department of Biological Sciences and Biotechnology, Botswana International University of Science and Technology, Palapye, Botswana
3. School of Biology and Environmental Sciences, University of Mpumalanga, Nelspruit 1200, South Africa
4. Kansas Biological Survey and the Biodiversity Institute, The University of Kansas, Lawrence, KS, USA
5. Department of Zoology and Entomology, Rhodes University, Makhanda, South Africa