1. Center of Deep Sea Research Institute of Oceanology, Chinese Academy of Sciences, Qingdao, P. R. China;
2. Center for Ocean Mega-Science, Chinese Academy of Sciences, Qingdao, P. R. China;
3. Key Laboratory of Marine Ecology and Environmental Sciences, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, P. R. China;
4. Laboratory for Marine Ecology and Environmental Science, Qingdao National Laboratory for Marine Science and Technology, Qingdao, China;
5. University of Chinese Academy of Sciences, Beijing, China