1. CAS Key Laboratory of Marine Ecology and Environmental Sciences, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China;
2. Laboratory for Marine Ecology and Environmental Science, Qingdao National Laboratory for Marine Science and Technology, Qingdao, China;
3. Center for Ocean Mega-Science, Chinese Academy of Sciences, Qingdao, China;
4. University of Chinese Academy of Sciences, Beijing, China