1. University of Coimbra, Centre for Functional Ecology-Science for People & the Planet, Department of Life Sciences, Calçada Martim de Freitas, 3000-456 Coimbra, Portugal
2. CESAM - Centre for Environmental and Marine Studies, Department of Biology, University of Aveiro, 3810-193 Aveiro, Portugal
3. Department of Life Sciences, Faculty of Sciences and Technology, University of Coimbra, Coimbra, Portugal
4. Department of Earth Sciences, Faculty of Sciences and Technology, University of Coimbra, Coimbra, Portugal