1. Graduate School of Advanced Integrated Studies in Human Survivability, Kyoto University , Kyoto, Japan
2. Division of Applied Life Sciences, Graduate School of Agriculture, Kyoto University , Kyoto, Japan
3. CREST, JST , Kyoto, Japan
4. Japan Society for the Promotion of Science , Kyoto, Japan
5. Department of Life Sciences, Graduate School of Bioresources, Mie University , Mie, Japan