1. School of Biology and Center of Excellence in Phylogeny of Living Organisms, College of Science, University of Tehran, Tehran, Iran
2. Department of Horticulture and Plant Protection, College of Agriculture, Shahrood University of Technology, Shahrood, Iran
3. Department of Biology, College of Sciences, Shiraz University, Shiraz, Iran
4. Staatliches Museum für Naturkunde Karlsruhe, Karlsruhe, Germany